1. National Institute for Agro-Environmental Sciences, 3-1-3 Kannondai, Tsukuba, 305-8604 Ibaraki, Japan.
2. Soil Microbiology Research Team, National Agricultural Research Center, Ibaraki 305-8666, Japan.
3. National Institute of Vegetable and Tea Science (NIVTS), National Agriculture and Food Research Organization (NARO), 3-1-1 Kannodai, Tsukuba 305-8666, Japan.
4. National Agriculture and Food Research Organization National Agricultural Research Center for Tohoku Region, Arai, Fukushima City, Fukushima 960-2156, Japan.
5. National Agricultural Research Center for Hokkaido Region, Shinsei, Memuro, Kasai, Hokkaido 082-0081, Japan.
6. Graduate School of Bio-Applications and Systems Engineering, Tokyo University of Agriculture and Technology, Tokyo 184-8588, Japan.
7. Field Science Center, Graduate School of Agriculture Science, Tohoku University, 232-3 Yomogida, Naruko-onsen, Osaki, Miyagi 989-6711, Japan.