1. Scientific Services, Ezemvelo KZN Wildlife, P.O. Box 13053, Cascades 3202, KwaZulu-Natal, South Africa
2. School of Animal, Plant and Environmental Sciences, University of the Witwatersrand, Private Bag X3, Johannesburg, WITS 2050, Gauteng, South Africa
3. Centre for Functional Biodiversity, School of Life Sciences, University of KwaZulu-Natal, Scottsville, South Africa