1. Department of Zoology and Faculty of Land and Food Systems, 6270 University Boulevard, University of British Columbia, Vancouver, BC V6T 1Z4, Canada.
2. IFM Biology, Division of Zoology, University of Linköping, SE-58183 Linköping, Sweden.
3. School of Biological Sciences, The University of Queensland, Brisbane, Queensland 4072, Australia.
4. University of Gothenburg, Department of Biological and Environmental Sciences, SE-405 30 Gothenburg, Sweden.