1. Department of Organismic and Evolutionary Biology and Harvard University Herbaria, Harvard University, Cambridge, MA 02138, USA.
2. Department of Plant Science, University of Pretoria, Private Bag X20, 0028 Hatfield, Pretoria, South Africa.
3. African Centre for DNA Barcoding, University of Johannesburg, APK Campus, PO Box 524, Auckland Park 2006, Johannesburg, South Africa.
4. Bolus Herbarium, Biological Sciences Department, University of Cape Town, Private Bag X3, Rondebosch 7700, South Africa; Department of Biology, Umaru Musa Yarádua University, Katsina, Nigeria.
5. Department of Geography, Environmental Management and Energy Studies, University of Johannesburg, APK Campus, PO Box 524, Auckland Park 2006, Johannesburg, South Africa.