Unusual preservation of the trace fossil Conostichus in middle Silurian carbonate facies of Indiana, USA

Abstract

The long-ranging, plug-shaped ichnofossil Conostichus, attributed to solitary polypoid cnidarians, is most commonly described in the Paleozoic from fine-grained siliciclastic sediments, with few descriptions from carbonate settings. The few described examples of putative Conostichus preserved in carbonate sedimentary rocks of Paleozoic age are essentially conical masses lacking recognizable external radial or longitudinal structures. Herein, we describe the occurrence of well-preserved examples of this ichnogenus within middle Silurian (Wenlock: Sheinwoodian) carbonate sedimentary rocks of the Massie Formation from southeastern Indiana, USA. These specimens represent isolated apical discs of Conostichus with prominent radiating physal impressions displaying duodecimal symmetry. Interestingly, well-preserved specimens co-occur with relatively poorly preserved (“typical carbonate”) specimens. The factors responsible for this unusual taphonomic state are unclear, but the most likely explanation is that exceptionally preserved specimens represent burrows that were somehow infilled, at least in their apical terminations, with fine-grained carbonate sediment, enhancing preservational fidelity. In contrast, other burrows had their apical regions passively filled with larger carbonate particles that could not preserve fine details. This occurrence indicates that early Paleozoic carbonate sediments are capable of preserving Conostichus, and potentially other ichnofossils, in similar modes to later Paleozoic siliciclastic deposits.