1. Royal British Columbia Museum, 675 Belleville Street, Victoria, BC V8W 9W2, Canada.
2. Department of Biology and School of Earth and Ocean Sciences, University of Victoria, P.O. Box 1700, Victoria, BC V8W 2Y2, Canada.
3. Department of Geography and the Environment, University of the Fraser Valley, 33844 King Road Abbotsford, BC V2S 7M8, Canada.
4. Department of Earth Sciences, The University of Western Ontario, London, ON N6A 5B7, Canada.