Description of a partial Dromiceiomimus (Dinosauria: Theropoda) skeleton with comments on the validity of the genus

Abstract

Dromiceiomimus brevitertius is a North American ornithomimid diagnosed primarily by the ratio of tibia length to femur length. It has recently, and perhaps incorrectly, been considered synonymous with Ornithomimus edmontonicus, with several authors questioning the utility of limb ratios in diagnosing taxa. While isolated ornithomimosaur material is common, specimens with sufficient diagnostic material to explore the question of synonymy are comparatively rare. The putative Dromiceiomimus specimen UALVP 16182 represents one of the few specimens in which diagnostic elements are available. It is therefore an important specimen for assessing the validity of Dromiceiomimus and for examining the utility of using limb proportions to diagnose ornithomimid taxa. In this paper, UALVP 16182 is described, the tibia/femur ratio is examined in closely related ornithomimid taxa, and the ratio is found to distinguish Dromiceiomimus from Gallimimus, Ornithomimus, and Struthiomimus. A phylogenetic analysis recovered Anserimimus and Ornithomimus as sister taxa with Dromiceiomimus as an outgroup. Comparison of the manus revealed differences in the morphology of metacarpal I and the flexor tubercle of manual ungual II-3. Differences also appear in the surangular and scapula. An examination of stratigraphic positions of various specimens indicates that Dromiceiomimus is generally higher in section than Ornithomimus, although there are too few specimens to be statistically significant. This study agrees with other studies in concluding that limb proportions are roughly isometric in small theropods like ornithomimids and that the tibia/femur ratio may therefore be useful for diagnosing certain small taxa. These findings suggest that Dromiceiomimus may indeed be a valid taxon.