1. 600 Cumberland Place, Nanaimo, BC V9T 4S5, Canada
2. Department of Earth, Environmental and Geographic Sciences, Irving K. Barber School of Arts and Sciences, University of British Columbia Okanagan, 1177 Research Road, Kelowna, BC V1V 1V7, Canada
3. Ministry of Forests, Lands and Natural Resource Operations and Rural Development, Wildlife and Habitat Branch, Suite 401-333 Victoria Street, Nelson, BC V1L 4K3, Canada