1. Department of Zoology and Entomology, University of Pretoria, Pretoria 0002, South Africa.
2. Sokoine University of Agriculture, P.O. Box 3000, Morogoro, Tanzania.
3. Mammal Research Institute, Department of Zoology and Entomology, University of Pretoria, Pretoria 0002, South Africa.
4. SARChI Chair of Mammal Behavioural Ecology and Physiology, Department of Zoology, University of Pretoria, Pretoria 0002, South Africa.