1. Department of Biological Science, University of Calgary, 2500 University Drive Northwest, Calgary, AB T2N 1N4, Canada.
2. Bamfield Marine Sciences Centre, 100 Pachena Road, Bamfield, BC V0R 1B0 Canada.
3. Department of Biological Sciences, University of Victoria, 3800 Finnerty Road, Victoria, BC V8P 4C2, Canada.