1. Environmental Science Program, Memorial University of Newfoundland, St. John’s, NL A1B 3X7, Canada.
2. School of Science and the Environment, Memorial University of Newfoundland, Corner Brook, NL A2H 5G4, Canada.
3. Institute of Agriculture, Natural Resources and Extension, University of Alaska Fairbanks, Fairbanks, AK 99775, USA.
4. Department of Natural Resources, McGill University, St Anne de Bellevue, QC H9X 3V9, Canada.