1. Faculty of Land and Food Systems, University of British Columbia, Vancouver, BC V6T 1Z4, Canada
2. BC Ministry of Forests, Kamloops, BC V2C 2T3, Canada
3. Faculty of Forestry, University of British Columbia, Vancouver, BC V6T 1Z4, Canada
4. Department of Botany, University of British Columbia, Vancouver, BC V6T 1Z4, Canada