1. Key Laboratory of Adaptation and Evolution of Plateau Biota, Northwest Institute of Plateau Biology, Chinese Academy of Sciences, No. 23, Xinning Road, Xining 810008, Qinghai, People’s Republic of China, Xining, Qinghai, China
2. University of Chinese Academy of Sciences, No. 19 (Jia), Shijingshan District, Beijing 100049, People’s Republic of China., Beijing, China;
3. State Key Laboratory of Cryosphere Science, Cold and Arid Regions Environmental and Engineering Research Institute, Chinese Academy of Sciences, Lanzhou, Gansu, China;
4. Key Laboratory of Adaptation and Evolution of Plateau Biota, Northwest Institute of Plateau Biology, Chinese Academy of Sciences, Xining, Qinghai, China
5. Key Laboratory of Crop Molecular Breeding of Qinghai Province, Northwest Institute of Plateau Biology, Chinese Academy of Sciences, Xining, Qinghai, China;
6. University of Chinese Academy of Sciences, Beijing, China;
7. Sichuan Agricultural University, Triticeae Research Institute, Huimin Road 211#, Gongping Town,Wenjiang, Chengdu City, Sichuan Province, Chengdu, China, 611130;
8. Northwest Institute of Plateau Biology Chinese Academy of Sciences, 34721, Xinning Road No.23, Xining, China, 810008, , ;