1. College of Life Sciences, Sichuan Agricultural University, Yaan, Sichuan 625014, China.
2. Lethbridge Research Centre, Agriculture and Agri-Food Canada, Lethbridge, AB T1 J 4B1, Canada.
3. Lacombe Research Centre, Agriculture and Agri-Food Canada 6000 C and E Trail, Lacombe, AB T4L 1W1, Canada.