1. Saint Mary's University, Biology, Halifax, Nova Scotia, Canada;
2. Agriculture and Agri-Food Canada, Dalhousie University, Faculty of Agriculture, PO Box 550, 100-5 Haley Institute, Truro, Nova Scotia, Canada, B2N 5E3, , ;
3. Saint Mary's University, Biology, 923 Robie Street, Halifax, Nova Scotia, Canada, B3H 3C3, , ;