1. Center for Post-doctoral Studies of Agricultural Resources and Environment, College of Land and Environment, Shenyang Agricultural University, Shenyang, Liaoning Province 98428, China
2. College of Forestry, Shenyang Agricultural University, Shenyang, Liaoning Province, China
3. Liaoning Academy of Forestry Sciences, Shenyang, Liaoning Province, China
4. State Key Laboratory of Tree Genetics and Breeding, Key Laboratory of Tree Breeding and Cultivation of State Forestry and Grassland Administration, Research Institute of Forestry, Chinese Academy of Forestry, Beijing 100091, China