Stand density biases the estimation of the site index especially on dry sites

Abstract

The site index (SI) has been widely used in forest management and silviculture. It relies on the assumption that the height of dominant trees in a stand is independent from the local density. However, research on climate change suggests that under certain moisture stress conditions, this may not hold. Here, based on 29 plots from five long-term research experiments, we tested the effect of local stand density on the SI of Norway spruce (Picea abies (L.) H. Karst). With generalized additive models (GAMM), we analyzed the effect of stand structure and climate predictors on SI. The two evaluated models revealed that local stand density and age had a significant effect on SI (p ≤ 0.001), showing a clear negative trend especially significant on sites with poor and dry soils, which may reduce the SI by a maximum of approximately 4 m for an increase in density of between 400 and 600 trees/ha. We stress that the physiological characteristics of Norway spruce, flat-rooting system and xeromorphism, especially when growing in pure stands, may explain these effects. Thus, density control and growth in mixtures may help to reduce the water stress and losses in height growth under future climate conditions.