1. Institute of Forestry and Conservation, John H. Daniels Faculty of Architecture, Landscape and Design, University of Toronto, Toronto, ON M5S 3B3, Canada
2. Great Lakes Forestry Centre, Canadian Forest Service, Natural Resources Canada, Sault Ste. Marie, ON P6A 2E5, Canada
3. Department of Statistical and Actuarial Sciences, University of Western Ontario, London, ON N6A 5B7, Canada