1. College of Forest Resources and Environmental Science, Michigan Technological University, Houghton, MI 49931, USA
2. School of Forest Science and Management, University of Alberta, Edmonton, AB T6G 2R3, Canada
3. Department of Forest Ecology and Management, Swedish University of Agricultural Sciences, Umeå 75007, Sweden
4. Scion (New Zealand Forest Research Institute), Rotorua, 3046, New Zealand