1. Department of Zoology and Entomology, University of Pretoria, Private Bag X20, Hatfield 0028, South Africa.
2. Wildlife and Reserve Management Research Group, Department of Zoology and Entomology, Rhodes University, PO Box 94, Grahamstown, 6140, South Africa.
3. SARChI Chair of Behavioural Ecology and Physiology, Department of Zoology and Entomology, University of Pretoria, Pretoria X20, Hatfield 0028, South Africa.