1. School of Resource and Environmental Management, Simon Fraser University, 8888 University Drive, Burnaby, BC V5A 1S6, Canada; Hakai Institute, P.O. Box 309, Heriot Bay, BC V0P 1H0, Canada.
2. B.C. Ministry of Forests, Lands and Natural Resource Operations, 103-2100 Labieux Rd., Nanaimo, BC V9T 6E9, Canada.