1. University of Jyvaskyla, Department of Biological and Environmental Science, P.O. Box 35, 40014 University of Jyvaskyla, Finland.
2. Simosol Oy, Rautatietori 4, 11130 Riihimäki, Finland.
3. Economics and Society, Natural Resources Institute Finland (Luke), P.O. Box 68, 80101 Joensuu, Finland.