1. Department of Ecology and Evolutionary Biology, University of Toronto 25 Harbord St. Toronto, ON M5S 3G5, Canada.
2. Pacific Forestry Centre, 506 Burnside Road West Victoria, BC V8Z 1M5, Canada.
3. Laurentian Forestry Centre. 1055 Du P.E.P.S. Street, P.O. Box 10380, Québec, QC G1V 4C7, Canada.