1. Center for Stock Assessment Research, University of California, Santa Cruz, Mail Stop SOE-2, Santa Cruz, CA 95064, USA.
2. Fisheries Ecology Division, Southwest Fisheries Science Center, National Marine Fisheries Service, National Oceanographic and Atmospheric Administration, 110 Shaffer Road, Santa Cruz, CA 95060, USA.
3. Department of Applied Mathematics and Statistics, Jack Baskin School of Engineering, University of California, Santa Cruz, Mail Stop SOE-2, Santa Cruz, CA 95064, USA.
4. Department of Biology, University of Bergen, Bergen, NO-5007, Norway.