1. Department of Biology, Memorial University of Newfoundland, St. John’s, NL A1B 3X9, Canada.
2. Department of Environment and Conservation, Government of Newfoundland and Labrador, St. John’s, NL A1B 4J6, Canada.
3. School for Resource and Environmental Studies, Dalhousie University, 6100 University Avenue, Halifax, NS B3H 3J5, Canada.