1. Environment and Climate change Canada, Science and Technology, 801-1550 ave d'Estimauville, Quebec City, Quebec, Canada, G1J 0C3, ;
2. Universite Laval, 4440, Centre d'étude de la forêt, Quebec, Quebec, Canada;
3. Environment and Climate change Canada, Canadian Wildlife Service, Quebec City, Quebec, Canada;
4. Carleton University, 13 Erinlea Court, Ottawa, Ontario, Canada, K2e 7c8, ;
5. Campus of McGill University, Department of Natural Resource Sciences Macdonald, 21,111 Lakeshore Road, Ste. Anne de Bellevue, Quebec, Canada, H9X 3V9, , ;