1. Pacific Salmon Ecology and Conservation Laboratory, Department of Forest and Conservation Sciences, University of British Columbia, Vancouver, BC V6T 1Z4, Canada
2. Pacific Salmon Foundation, Vancouver, BC V6J 4S6, Canada
3. Department of Ecology and Evolutionary Biology, University of Toronto, Toronto, ON M5S 3B2, Canada
4. Institute of Ocean Sciences, Fisheries and Oceans Canada, Sidney, BC V8L 5T5, Canada
5. Fisheries Science Department, Columbia River Inter-Tribal Fish Commission, Portland, OR 97232, USA
6. Pacific Biological Station, Fisheries and Oceans Canada, Nanaimo, BC, Canada
7. Institute for the Oceans and Fisheries, University of British Columbia, Vancouver, BC V9T 6N7, Canada
8. Fisheries and Oceans Canada, School of Resource and Environmental Management, Simon Fraser University, Science Branch, Burnaby, BC V5A 1S6, Canada