1. Southwest University for Nationalities, 66336, College of Life Science and Technology, Sichuan, China;
2. Southwest University for Nationalities, Chengdu, China;
3. Southwest University for Nationalities, College of Life Science and Technology, 16# South section 4th, First Ring Road, Chengdu, Sichuan, China, 610041, , ;
4. Key Laboratory of Sichuan Province for Qinghai-Tibetan Plateau Animal Genetic Resource Reservation and Exploitation, Institution of Qinghai-Tibetan Plateau, 16#, Yihuan Road, Chengdu City, Sichuan Province, China, Chengdu, China, 610041
5. Canada;