1. 301 Pebble Creek Dr., Garland, TX 75040
2. Key Laboratory of Vertebrate Evolution and Systematics, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing, China 100044
3. CAS-Center of Excellence in Life and Paleoenvironment, Beijing, China 100044
4. University of Chinese Academy of Sciences, Beijing, China 100039