1. Avian Zoonosis Research Center, Faculty of Agriculture, Tottori University, Tottori, Japan
2. Department of Joint Veterinary Medicine, Faculty of Agriculture, Tottori University, Tottori, Japan
3. Laboratory of Microbiology, Department of Disease Control, Faculty of Veterinary Medicine, Hokkaido University, Hokkaido, Japan
4. Division of Global Epidemiology, International Institute for Zoonosis Control, Hokkaido University, Hokkaido, Japan
5. International Collaboration Unit, International Institute for Zoonosis Control, Hokkaido University, Hokkaido, Japan