1. Laboratory of Zoonotic Diseases, Faculty of Applied Biological Sciences, Gifu University, 1–1 Yanagido, Gifu, Gifu 501–1193, Japan
2. The United Graduate School of Veterinary Sciences, Gifu University, 1–1 Yanagido, Gifu, Gifu 501–1193, Japan
3. Department of Biochemistry and Molecular Biology, Bio21 Molecular Science and Biotechnology Institute, 30 Flemington Rd., The University of Melbourne, Victoria 3010, Australia