1. Academy of Animal and Veterinary Medicine, University of Qinghai, No. 1, Weier road, Sci-biological Industry Areas, Xining, Qinghai, the People of Republic of China
2. Laboratory of Veterinary Parasitic Diseases, Interdisciplinary Graduate School of Medicine and Veterinary Medicine, University of Miyazaki, 1–1 Gakuen-Kibanadai-Nishi, Miyazaki 889–2192, Japan
3. Institute of Livestock and Grassland Science, National Agriculture and Food Research Organization (NARO), 375–716 Shiono, Miyota, Nagano 389–0201, Japan
4. Laboratory of Veterinary Anatomy, Department of Veterinary Sciences, Faculty of Agriculture, University of Miyazaki, 1–1 Gakuen-Kibanadai-Nishi, Miyazaki 889–2192, Japan
5. Laboratory of Veterinary Parasitic Diseases, Department of Veterinary Sciences, Faculty of Agriculture, University of Miyazaki, 1–1 Gakuen-Kibanadai-Nishi, Miyazaki 889–2192, Japan