1. Laboratory of Veterinary Parasitic Diseases, Interdisciplinary Graduate School of Medicine and Veterinary Medicine, University of Miyazaki, 1–1 Gakuen-Kibanadai-Nishi, Miyazaki, 889–2192, Japan
2. Academy of Animal and Veterinary Medicine, University of Qinghai, No. 1, Weier road, Sci-biological Industry Areas, Xining, Qinghai, the People of Republic of China
3. Graduate School of Agriculture, University of Miyazaki, 1–1 Gakuen-Kibanadai-Nishi, Miyazaki, 889–2192, Japan
4. Laboratory of Veterinary Anatomy, Department of Veterinary Sciences, Faculty of Agriculture, University of Miyazaki, 1–1 Gakuen-Kibanadai-Nishi, Miyazaki, 889–2192, Japan
5. Division for International Cooperation and Education, Center for Animal Disease Control, University of Miyazaki, 1–1 Gakuen-Kibanadai-Nishi, Miyazaki, 889–2192, Japan