1. Laboratory of Small Animal Internal Medicine, Joint Faculty of Veterinary Medicine, Kagoshima University, 1–21–24 Korimoto, Kagoshima, Kagoshima 890–0065, Japan
2. Laboratory of Parasitology, Joint Faculty of Veterinary Medicine, Kagoshima University, 1–21–24 Korimoto, Kagoshima, Kagoshima 890–0065, Japan
3. Laboratory of Ecology and Systematics, Faculty of Science, University of the Ryukyus, 1 Senbaru, Nishihara, Okinawa 903–0213, Japan
4. Iriomote Wildlife Conservation Center, Ministry of the Environment, Komi, Taketomi, Yaeyama-gun, Okinawa 907–1432, Japan
5. Veterinary Teaching Hospital, Kagoshima University, 1–21–24 Korimoto, Kagoshima, Kagoshima 890–0065, Japan