1. Laboratory of Infectious Disease, School of Veterinary Medicine, Azabu University, 1-17-71 Fuchinobe, Chuo-ku, Sagamihara, Kanagawa 252-5201, Japan
2. Research and Education Center for Prevention of Global Infectious Diseases of Animals, Tokyo University of Agriculture and Technology, 3-5-8 Saiwai, Fuchu, Tokyo 183–8509, Japan
3. The United Graduate School of Veterinary Sciences, Gifu University, Yanagito, Gifu 501-1193, Japan
4. Faculty of Veterinary Science, Nippon Veterinary and Life Science University, Musashino-shi, Tokyo 180-8602, Japan
5. Value Farm Consulting Co., Ltd., 1704-3 Nishi Oi, Tsukuba, Ibaraki 300-1260, Japan
6. Drexel University Dornsife School of Public Health, Philadelphia PA 19104, USA
7. Centre for Infection and Immunity, Mailman School of Public Health, Columbia University, New York, NY 10032, USA