1. Ecological Genetics Analysis Section, Center for Environmental Biology and Ecosystem Studies, National Institute for Environmental Studies, 16–2 Onogawa, Tsukuba, Ibaraki 305–8506, Japan
2. Yambaru Wildlife Conservation Center, 263–1 Hiji, Kunigami, Okinawa 905–1413, Japan
3. Conservation & Animal Welfare Trust, 308–7–205 Maehara, Uruma, Okinawa 904–2235, Japan