1. Novosibirsk State University, Novosibirsk, 630090, Russia
2. Research Institute of Experimental and Clinical Medicine, Novosibirsk, 630117, Russia
3. Laboratory of Microbiology, Department of Disease Control, Graduate School of Veterinary Medicine, Hokkaido University, Sapporo, Hokkaido 060-0818, Japan
4. Global Station for Zoonosis Control, Global Institution for Collaborative Research and Education, Hokkaido University, Sapporo, Hokkaido 001-0020, Japan
5. School of Biomedicine, Far Eastern Federal University, Vladivostok, 690950, Russia
6. Institute of Biology and Soil Science, Far Eastern Branch of Russian Academy of Sciences, Vladivostok, 690022, Russia