1. Division of Collaboration and Education, Hokkaido University Research Center for Zoonosis Control, Kita 20 Nishi 10, Kita-ku, Sapporo, Hokkaido 001-0020 Japan
2. Hokudai Center for Zoonosis Control in Zambia, the University of Zambia, Lusaka, 10101 Zambia
3. Graduate School of Science and Technology, Niigata University, Ikarashi 2 no-cho 8050, Nishi-ku, Niigata, Niigata 950-2181 Japan
4. Unit of Risk Analysis and Management, Hokkaido University Research Center for Zoonosis Control, Kita 20 Nishi 10, Kita-ku, Sapporo, Hokkaido 001-0020 Japan
5. Laboratory of Parasitology, Faculty of Veterinary Medicine, Graduate School of Infectious Diseases, Hokkaido University, Kita 18 Nishi 9, Kita-ku, Sapporo, Hokkaido 060-0818 Japan
6. Global Institution for Collaborative Research and Education (GI-CoRE), Hokkaido University, Kita 20 Nishi 10, Kita-ku, Sapporo, Hokkaido 001-0020 Japan