1. Department of Biomedicine, Graduate School of Medical Sciences, Kyushu University, 3-1-1 Maidashi Higashi-ku, Fukuoka 812-8582, Japan
2. Center of Biomedical Research, Research Center for Human Disease Modeling, Graduate School of Medical Sciences, Kyushu University, 3-1-1 Maidashi Higashi-ku, Fukuoka 812-8582, Japan
3. Joint Faculty of Veterinary Medicine, Transboundary Animal Diseases Research Center, Kagoshima University, 1-21-24 Kagoshima-shi, Kagoshima 890-0065, Japan
4. Biotechnology Development Laboratories, Health Care Solutions Research Institute, Kaneka Corporation, 1-8 Takasago-shi, Hyogo 676-8688, Japan