1. Graduate School of Life and Environmental Sciences, Kyoto Prefectural University, Shimogamo Sakyo-ku, Kyoto 606–8522, Japan
2. Transdisciplinary Research Integration Center, National Institute of Polar Research, 10–3 Midori-cho, Tachikawa, Tokyo 190–8518, Japan
3. Transdisciplinary Research Integration Center, 4–3–13 Toranomon, Minato-ku, Tokyo 105–0001, Japan
4. College of Bioresource Science, Nihon University, Fujisawa 252–0880, Japan