1. Laboratory of Wildlife Biology and Medicine, Department of Environmental Veterinary Sciences, Faculty of Veterinary Medicine, Hokkaido University, Hokkaido, Japan
2. Division of International Research Promotion, International Institute for Zoonosis Control, Hokkaido University, Hokkaido, Japan
3. Department of Animal Medicine, Faculty of Veterinary Medicine, South Valley University, Qena, Egypt
4. Laboratory of Parasitology, Department of Disease Control, Faculty of Veterinary Medicine, Hokkaido University, Hokkaido, Japan
5. Biodiversity Division, National Institute for Environmental Studies, Ibaraki, Japan
6. Institute of Biodiversity and Environmental Conservation, Universiti Malaysia Sarawak, Sarawak, Malaysia