1. School of Biology and Herbarium USJ, University of Costa Rica, 11501-2060 San Pedro de Montes de Oca, San José, Costa Rica.
2. Herbarium CR, National Museum of Costa Rica, P.O. Box 749-1000, San José, Costa Rica.
3. Department of Botany, Birge Hall, University of Wisconsin, Madison, Wisconsin 53706, U.S.A.