1. a Centre for Ecology and Hydrology , Wallingford, Oxfordshire, OX10 8BB, UK E-mail:
2. b Department of Geography , Loughborough University , Loughborough, UK
3. c Alconbury Consultants , Alconbury, UK
4. d Hull International Fisheries Institute, University of Hull , Hull, UK
5. e Environment Agency , Spalding, UK
6. f Environment Agency , Leeds, UK
7. g Department of Biological Sciences , University of Cape Town , South Africa
8. h Department of Geography , University of Dundee , Dundee, UK
9. i Scottish Environment Protection Agency , Stirling, UK