1. Lung and Allergy Research Centre, Diamantina Institute, The University of Queensland, Brisbane, Queensland, Australia;
2. Priority Research Centre for Healthy Lungs, The University of Newcastle, New Lambton Heights, New South Wales, Australia;
3. School of Biomedical Sciences and Pharmacy and
4. Immune Health Research Program, Hunter Medical Research Institute, New Lambton Heights, New South Wales, Australia
5. School of Medicine and Public Health, The University of Newcastle, Callaghan, New South Wales, Australia; and