1. Graduate School of Animal Bioscience, IPB University, Kampus IPB Dramaga, Bogor 16680, Indonesia
2. Research Center for Biosystematics and Evolution, National Research and Innovation Agency (BRIN), Government of Indonesia, Cibinong 16911, Indonesia
3. Department of Biology, Faculty of Mathematics and Natural Sciences, IPB University, Kampus IPB Dramaga, Bogor 16680, Indonesia
4. Centre de Recherche sur la Biodiversité et l'Environnement, UMR 5300, CNRS, IRD, Université Paul Sabatier, Bâtiment 4R1, 118 Route de Narbonne, 31062 cedex 9 Toulouse, France
5. Research Centre for Applied Zoology, National Research and Innovation Agency (BRIN), Government of Indonesia, Cibinong 16911, Indonesia