1. Sam Noble Oklahoma Museum of Natural History and Department of Biology, University of Oklahoma, 2401 Chautauqua Avenue, Norman, OK 73072–7029, USA
2. Biodiversity Institute and Department of Ecology and Evolutionary Biology; University of Kansas, 1345 Jayhawk Boulevard, Lawrence, KS 66045–7593, USA
3. Department of Biology, National Museum of Natural Science, Taichung 404, Taiwan