1. a Department of Plant Science , McGill University , 21111 Lakeshore Road, Ste-Anne-de-Bellevue, Quebec , H9X 3V9 , Canada
2. b Department of Crop and Soil Science , Oregon State University , Corvallis , OR , 97331 , USA
3. c Scottish Crop Research Institute, Invergowrie , Dundee , DD2 5DA , Scotland, UK