1. Faculty of Veterinary Science, University of Melbourne, Parkville, Victoria 3010, Australia
2. Faculty of Veterinary Science, University of Melbourne, Veterinary Clinical Centre, Werribee, Victoria 3030, Australia
3. School of Earth & Environmental Sciences, University of Adelaide, Adelaide, South Australia 5005, Australia
4. Conservation Ark, Zoos South Australia, Adelaide, South Australia 5000, Australia
5. These authors contributed equally