1. Canadian Wildlife Health Cooperative, Department of Pathology and Microbiology, Atlantic Veterinary College, University of Prince Edward Island, 550 University Avenue, Charlottetown, Prince Edward Island C1A 4P3, Canada
2. Environment and Climate Change Canada, Canadian Wildlife Service, 45 Alderney Drive, Dartmouth, Nova Scotia B2Y 2N6, Canada
3. Department of Biology, Acadia University, 33 Westwood Avenue, Wolfville, Nova Scotia B4P 2R6, Canada
4. Sable Island Institute, PO Box 11, Halifax Central, Halifax, Nova Scotia B3J 2L4, Canada