1. Canadian Wildlife Health Cooperative, Western College of Veterinary Medicine, University of Saskatchewan, 52 Campus Drive, Saskatoon, S7N 5B4 Saskatchewan, Canada
2. Epidemiology of Zoonoses and Public Health Research Group (GREZOSP), Faculty of Veterinary Medicine, Université de Montréal, 3200, rue Sicotte, Saint-Hyacinthe (Québec), J2S 2M2 Quebec, Canada
3. Department of Veterinary Pathology, Western College of Veterinary Medicine, University of Saskatchewan, 52 Campus Drive, Saskatoon, S7N 5B4 Saskatchewan, Canada
4. Department of Viroscience, Erasmus University Medical Centre, PO Box 2040, 3000 CA Rotterdam, the Netherlands